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In order that primeval men, or the ape-like progenitors of man, should become social, they must have acquired the same instinctive feelings, which impel other animals to live in a body… They would have felt uneasy when separated from their comrades, for whom they would have felt some degree of love; they would have warned each other of danger, and have given mutual aid in attack or defence. All this implies some degree of sympathy, fidelity and courage. Such social qualities… were no doubt acquired… through natural selection, aided by inherited habit. When two tribes… came into competition, if… the one tribe included a great number of courageous, sympathetic and faithful members, who were always ready to warn each other of danger, to aid and defend each other, this tribe would succeed better and conquer the other.
Charles DARWIN, Descent of Man, Chapter 5